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They compiled a 12–8–1 record in these seasons, playing as an independent from 1887–1890.
In 1891, the Pennsylvania Intercollegiate Football Association (PIFA) was formed.
The team is also #8 all-time in total-wins, one game behind Oklahoma and Alabama.
Bill Hollenback took over the Nittany Lions as head coach for the 1909 season and went undefeated at 5–0–2, Paterno spent 46 seasons as the head football coach, the longest tenure of any head coach in the FBS, and 16 more seasons as an assistant, making his 62 total years coaching at Penn State the most of any coach at any school.
Following the win, Penn State was ranked 24th in the AP Poll, its first ranking since 2011.